Frasier returns from Acapulco after a torrid affair with a supermodel/zoologist who's sworn him to secrecy regarding her identity. No one believes Frasier when he finally divulges her identity, instead believing that Frasier's finally lost his marbles.
Frasier is delighted to be a handpicked celebrity on a ""luxury"" cruise until he discovers it's luxurious in name only. Niles and Maris are on the outs after she tells him she'll be vacationing in Switzerland on their anniversary.
Frasier tags along with Daphne to her favorite pub and realizes he's been missing a Cheers-like atmosphere ever since he left Boston. They find themselves at odds with one another when Daphne insists that Frasier stop frequenting ""her"" place.
Niles and Maris' new counselor, Dr. Schenkman, advises Niles to inject some spontaneity into their relationship, inspiring Niles to surprise Maris in her bedroom. Niles isn't quite prepared for what he finds.
The Cranes are reunited with a branch of the family when Frasier's cousin, Nikos, is getting married. First, Frasier has to soothe old wounds with Martin's sister-in-law, incurred when Frasier urged Nikos to pursue a career in juggling rather than medicine.
The whole radio station is buzzing over the incredibly attractive and talented new radio host, Dr. Clint Webber. Frasier tries desperately to find Clint's "Achilles heel," while Martin is thrown out of the only store in Seattle selling Eddie's favorite dog food.
Frasier refuses to comply when the new station manager mandates that on-air talent must read commercials during their programs. Frasier leads his forces to ""The Black Tower"" to protest, reckoning that the might of the on-air talent will force management to accede to their demands.